Under review
Morais, A.R.; Hepp, F.; Pansonato, A.; Santos, C.E.; Haddad, C.F.B.; Strüssmann, C.; Nascimento, L.B.; Toledo, L.F.; Duarte, M.H.L.; Wachlevski, M.; Simões, P.I.; Bastos, R.P.; Carvalho-e-Silva, S.P.; Dena, S.; Carvalho, T.R.; Forti, L.R. Amphibian sound recordings in Brazil are geographically and taxonomically biased and cover less than two-thirds of native species.
Cassini, C.S.; Carvalho, T.R.; Taucce, P.P.G.; Haddad, C.F.B.; Solé, M. Unexplored urban diversity: a new species of Adenomera (Anura, Leptodactylidae) related to A. ajurauna from the Atlantic Forest of southeastern and southern Brazil.
Zaracho, V.H.; Lavilla, E.O.; Carvalho, T.R.; Motte, M.; Basso, N.G. In pursuit of a nominal anuran species: redescription of Adenomera diptyx (Boettger, 1885) (Leptodactylidae) and description of a closely related new species.
Marinho, P.M.; Santos, M.T.T.; Faivovich, J., Lyra, M.L.; Giaretta, A.A.; Haddad, C.F.B.; Carvalho, T.R. A new species of the Aplastodiscus albosignatus group (Hylinae: Cophomantini) from the northern Mantiqueira Mountain Range.
Accepted for publication
Bang, D.L.; Bockmann, F.A.; Giaretta, A.A.; Carvalho, T.R. The vocal repertoire of two Bokermannohyla (Anura: Hylidae) species, with an overview of the advertisement call diversity in the genus. Journal of Herpetology.
Complete list of publications
2022
Santos, M.T.T.; Barata, I.M.; Ferreira, R.B.; Haddad, C.F.B.; Gridi-Papp, M.; Carvalho, T.R. Complex acoustic signals in Crossodactylodes (Leptodactylidae, Paratelmatobiinae): a frog genus historically regarded as voiceless. Bioacoustics, 31: 175-190. (PDF)
Carvalho, T.R.; Fouquet, A.; Lyra, M.L.; Giaretta, A.A.; Costa-Campos, C.E.; Rodrigues, M.T.; Haddad, C.F.B.; Ron, S.R. Species diversity and systematics of the Leptodactylus melanonotus group (Anura, Leptodactylidae): review of diagnostic traits and a new species from the Eastern Guiana Shield. Systematics and Biodiversity, 20(1): #2089269.
Novaes-e-Fagundes, G.; Lyra, M.L.; Loredam, V.; Carvalho, T.R.; Haddad, C.F.B.; Rodrigues, M.T.; Baldo, D.; Barrasso, D.; Loebmann, D.; Ávila, R.W.; Brusquetti, F.; Prudente, A.L.C.; Wheeler, W.C.; Orrico, V.G.D.; Peloso, P.L.V. A tale of two bellies: systematics of the oval frogs (Anura: Microhylidae: Elachistocleis Parker, 1927). Zoological Journal of the Linnean Society, online first: 1-24.
Taucce, P.P.G.; Costa-Campos, C.E.; Carvalho, T.R.; Michalski, F. Anurans (Amphibia: Anura) of the Brazilian state of Amapá, eastern Amazonia: species diversity and knowledge gaps. European Journal of Taxonomy, 836: 96-130.
Fonseca, W.L.; Almeida, M.R.N.; Nascimento, J.A.F.; Moraes, L.G.L.; Machado, R.A.; Bernarde, P.S.; Costa-Campos, C.E.; Carvalho, T.R. The advertisement call of Allophryne resplendens (Anura: Allophrynidae) from southwestern Brazilian Amazonia. Zootaxa, 3158: 196-200.
2021
Carvalho, T.R.; Seger, K.R.; Magalhães, F.M.; Lourenço, L.B.; Haddad, C.F.B. Systematics and cryptic diversification of Leptodactylus frogs in the Brazilian campo rupestre. Zoologica Scripta, 50(3): 300-317.
Carvalho, T.R.; Moraes, L.C.J.L.; Lima, A.P.; Fouquet, A.; Peloso, P.L.V.; Pavan, D.; Drummond, L.O.; Rodrigues, M.T.; Giaretta, A.A.; Gordo, M.; Neckel-Oliveira, S.; Haddad, C.F.B. Systematics and historical biogeography of Neotropical foam-nesting frogs of the Adenomera heyeri clade (Leptodactylidae), with the description of six new Amazonian species. Zoological Journal of the Linnean Society, 191(2): 395-433. (PDF)
Gazoni, T.*; Lyra, M.L.*; Ron, S.R.; Strüssmann, C.; Baldo, D.; Narimatsu, H.; Pansonato, A.; Schneider R.G.; Giaretta, A.A.; Haddad, Parise-Maltempi, P.P.; Carvalho, T.R*. Revisiting the systematics of the Leptodactylus melanonotus group (Anura: Leptodactylidae): redescription of L. petersii and revalidation of its junior synonyms. Zoologischer Anzeiger, 290: 117-134. (request by e-mail)
* Equal authorship.
Fouquet, A.*; Marinho, P.*; Réjaud, A.; Carvalho, T.R.; Caminer, M.; Jansen, M.; Rainha, R.N.; Rodrigues, M.T.; Werneck, F.P.; Lima, A.P.; Hrbek, T.; Giaretta, A.A.; Venegas, P.J.; Chávez, G; Ron, S. Systematics and biogeography of the Boana albopunctata species group, with the description of two new species from Amazonia. Systematics and Biodiversity, 19(4): 375-399. (request by e-mail)
* Shared co-first authorship.
Carvalho, T.R.; Seger, K.R.; Magalhães, F.M.; Lourenço, L.B.; Haddad, C.F.B. On the availability of the recently proposed nomen Leptodactylus avivoca (Anura, Leptodactylidae). Bionomina, 22(1): 42-45. (request by e-mail)
2020
Carvalho, T.R.; Giaretta, A.A. Diversidade, distribuição e conservação dos anfíbios anuros. In: Vasconcelos, H.L. (Ed.). Ecologia e conservação dos cerrados, campos e florestas do Triângulo Mineiro e sudeste de Goiás. Editora Regência e Arte. (PDF)
Magalhães, F.M.; Lyra, M.L.; Carvalho, T.R.; Baldo, D.; Brusquetti, F.; Burella, P.; Colli, G.R.; Gehara, M.C.; Giaretta, A.A.; Haddad, C.F.B.; Langone, J.A.; López, J.A.; Napoli, M.F.; Santana, D.J.; de Sá, R.O.; Garda, A.A. Taxonomic review of South American butter frogs: phylogeny, biogeographic patterns, and species delimitation in the Leptodactylus latrans species group (Anura: Leptodactylidae). Herpetological Monographs, 34: 131-177. (request by e-mail)
Carvalho, T.R.; Simões, P.I.; Gagliardi-Urrutia, G.; Rojas-Runjaic, F.J.M.; Haddad, C.F.B.; Castroviejo-Fisher, S. A new forest-dwelling frog species of the genus Adenomera (Leptodactylidae) from northwestern Brazilian Amazonia. Copeia, 108(4): 924-937. (request by e-mail)
Taucce, P.P.G.; Costa-Campos, C.E.; Haddad, C.F.B.; Carvalho, T.R. A new Amazonian species of the diminutive frog genus Adelophryne (Anura: Brachycephaloidea: Eleutherodactylidae) from the state of Amapá, northern Brazil. Copeia, 108(4): 746-757. (request by e-mail)
Cassini, C.S.; Taucce, P.P.G.; Carvalho, T.R.; Fouquet, A.; Solé, M.; Haddad, C.F.B.; Garcia, P.C.A. One step beyond a broad molecular phylogenetic analysis: species delimitation of Adenomera marmorata Steindachner, 1867 (Anura: Leptodactylidae). Plos One, 15(2): e0229324. (PDF)
Silva, L.A.; Magalhães, F.M.; Thomassen, H.; Leite, F.S.F.; Garda, A.A.; Brandão, R.A.; Haddad, C.F.B.; Giaretta, A.A.; Carvalho, T.R. Unraveling the species diversity and relationships in the Leptodactylus mystaceus complex (Anura: Leptodactylidae), with the description of three new Brazilian species. Zootaxa, 4779(2): 151-189. (request by e-mail)
Marinho, P.; Costa-Campos, C.E.; Pezzuti, T.L.; Dias-Souza, M.R.; Haddad, C.F.B.; Giaretta, A.A.; Carvalho, T.R. The Amapá tree frog Boana dentei (Bokermann, 1967): diagnosis and redescription (Hylinae: Cophomantini). Journal of Natural History, 54(15-16): 971-990. (request by e-mail)
Carvalho, T.R.*; Moraes, L.J.C.L.*; Angulo, A.; Werneck, F.P.; Icochea, J.; Lima, A.P. New acoustic and molecular data shed light on the poorly known Amazonian frog Adenomera simonstuarti
(Leptodactylidae): implications for distribution and conservation. European Journal of Taxonomy, 682: 1-18. (PDF)
* Shared co-first authorship.
Bang, D.L.; Lisboa, B.S.; Teixeira, B.F.V.; Giaretta, A.A.; Carvalho, T.R. A comparative acoustic analysis of species of Vitreorana (Anura: Centrolenidae) from the Brazilian Atlantic Forest and Cerrado, with a description of the call of V. baliomma and insights into the taxonomic status of Cerrado populations. Phyllomedusa, 19(1): 35-47. (PDF)
Andrade, F.S.; Haga, I.A.; Lyra, M.L.; Carvalho, T.R.; Haddad, C.F.B.; Giaretta, A.A.; Toledo, L.F. Reassessment of the taxonomic status of Pseudopaludicola parnaiba (Anura, Leptodactylidae, Leiuperinae), with the description of a new cryptic species from the Brazilian Cerrado. European Journal of Taxonomy, 679: 1-36. (PDF)
Carvalho, T.R.; Angulo, A.; Barrera, D.A.; Aguilar-Puntriano, C.; Haddad, C.F.B. Hiding in plain sight: a fourth new cryptic species of the Adenomera andreae clade (Anura: Leptodactylidae) from southwestern Amazonia. Herpetologica, 76(3): 304-314. (request by e-mail)
Carvalho, T.R.; Roberto, I.J.; Dias, E.M.; Santos, R.L.; Santos, E.M. The advertisement call of Physalaemus caete (Leptodactylidae), an endangered anuran species endemic to Brazil's northern Atlantic Forest. Zootaxa, 4822(3): 439-442. (request by e-mail)
2019
Carvalho, T.R.; Giaretta, A.A.; Maciel, N.M.; Barrera, D.A.; Aguilar-Puntriano, C.; Haddad, C.F.B.; Kokubum, M.N.C.; Menin, M.; Angulo, A. On the uncertain taxonomic identity of Adenomera hylaedactyla (Cope, 1868) and the composite type series of A. andreae (Müller, 1923) (Anura, Leptodactylidae). Copeia, 107(4): 708-723. (request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Angulo, A.; Kokubum, M.N.C.; Barrera, D.A.; Souza, M.B.; Haddad, C.F.B.; Giaretta, A.A. A new cryptic species of the Adenomera andreae clade from southwestern Amazonia (Anura, Leptodactylidae). Herpetologica, 75(3): 233-246. (request pdf at thiago_decarvalho@yahoo.com.br)
Marinho, P.; Costa-Campos, C.E.; Sanches, P.R.; Carvalho, T.R. The release call of Rhaebo guttatus (Anura: Bufonidae). Studies on Neotropical Fauna and Environment, online first.
(request pdf at thiago_decarvalho@yahoo.com.br)
Santos, M.T.T.; Oliveira, S.H.; Carvalho, T.R.; Zaidan, B.F.; Silva, N.R.; Berneck, B.V.M.; Garcia, P.C.A. A new species of Paratelmatobius (Anura: Leptodactylidae: Paratelmatobiinae) from the Atlantic Forest of southern Brazil. Zootaxa, 4648(3): 473-493. (request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Haddad, C.F.B.; Gridi-Papp, M. Tonal calls as a bioacoustic novelty in two Atlantic Forest species of Physalaemus (Anura: Leptodactylidae). Acta Herpetologica, 14(1): 21-26. (PDF)
Carvalho, T.R.; Cassini, C.S.; Taucce, P.P.G.; Haddad, C.F.B. A new, morphologically cryptic species of Adenomera closely related to A. araucaria from the Atlantic Forest of southern Brazil (Anura, Leptodactylidae). Journal of Herpetology, 53(2): 131-143.
(request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Giaretta, A.A.; Angulo, A.; Haddad, C.F.B.; Peloso, P.L.V. A new Amazonian species of Adenomera (Anura: Leptodactylidae) from the Brazilian state of Pará: a tody-tyrant voice in a frog. American Museum Novitates, 3919: 1-21. (PDF)
Roberto, I.J.; Santos, E.M.; Carvalho, T.R. The vocalization of Gastrotheca fissipes (Boulenger, 1888) (Anura, Hemiphractidae) from the state of Pernambuco, northeastern Brazil. Zootaxa, 4543(2): 284-286.
(request pdf at thiago_decarvalho@yahoo.com.br)
2018
Andrade, F.S.; Haga, I.A.; Lyra, M.L.; Carvalho, T.R.; Haddad, C.F.B., Giaretta, A.A., Toledo, L.F. A new species of Pseudopaludicola (Anura: Leptodactylidae: Leiuperinae) from the state of Minas Gerais, Brazil. European Journal of Taxonomy, 480: 1-25. (PDF)
Marinho, P.; Carvalho, T.R.; Bang, D.L.; Teixeira, B.F.V.; Azarak, P.A.; Costa-Campos, C.E. & Giaretta, A.A. Advertisement calls, intraspecific variation and species diagnosis of six Brazilian species of Elachistocleis (Anura: Microhylidae: Gastrophryninae). Zootaxa, 4521(3): 357-375.
(request pdf at thiago_decarvalho@yahoo.com.br)
Costa-Campos, C.E.; Carvalho, T.R. The advertisement call of the Hoogmoed's harlequin toad Atelopus hoogmoedi Lescure, 1974 from northern Brazil (Anura, Bufonidae). Zootaxa, 4521(1): 141-144.
(request pdf at thiago_decarvalho@yahoo.com.br)
Andrade, F.S.; Haga, I.A.; Carvalho, T.R.; Martins, L.B.; Giaretta, A.A. An acoustic and morphometric evaluation of the geographic distribution of Phyllomedusa burmeisteri (Anura: Phyllomedusidae), with comments on P. bahiana. Phyllomedusa, 17(1): 39-59. (PDF)
Vidigal, I.; Carvalho, T.R.; Clemente-Carvalho, R.B.G. & Giaretta, A.A. Vocalizations, tadpole, and natural history of Crossodactylus werneri Pimenta, Cruz & Caramaschi, 2014 (Anura: Hylodidae), with comments on distribution and intraspecific variation. Zootaxa, 4388(1): 061-075.
(request pdf at thiago_decarvalho@yahoo.com.br)
2017
Carvalho, T.R.; Azarak, P.A.; Bang, D.L.; Duellman, W.E. & Giaretta, A.A. A reassessment of the vocalization and distribution of Scinax exiguus (Duellman, 1986) (Anura: Hylidae) in the Amazonian savanna of Roraima, northern Brazil, with the description of its aggressive call. Neotropical Biodiversity, 3(1): 196-202. (PDF)
Carvalho, T.R.; Bang, D.L.; Teixeira, B.F.V. & Giaretta, A.A. First record of Boana alfaroi (Caminer & Ron, 2014) (Anura: Hylidae) in Brazil. Check List, 13(4): 135-139. (PDF)
Haga, I.A.; Carvalho, T.R.; Andrade, F.S. & Giaretta, A.A. Advertisement and aggressive calls of Pithecopus azureus (Anura: Phyllomedusidae) from border of Brazil and Paraguay . Phyllomedusa, 16(1): 47-56. (PDF)
Andrade, F.S.; Carvalho, T.R.; Martins, L.B. & Giaretta, A.A. Reassessment of the vocal repertoire of a nest-building gladiator frog, Boana pardalis (Anura, Hylidae, Cophomantinae): implications for its diagnosis within the B. faber species group. Studies on Neotropical Fauna and Environment, 52(1): 1-11. (PDF)
Bang, D.L.; Carvalho, T.R.; Andrade, F.S.; Aquemi, I.H.; Bernardes, C.S. & Giaretta, A.A. Vocalization of Scinax haddadorum (Anura: Hylidae), with further notes on the vocalization of the morphologically similar S. rupestris. Neotropical Biodiversity, 3(1): 117-124. (PDF)
Andrade, F.S.; Leite, F.S.; Carvalho, T.R.; Bernardes, C.S. & Giaretta, A.A. First record of Pseudopaludicola pocoto Magalhães, Loebmann, Kokubum, Haddad & Garda, 2014 (Anura, Leptodactylidae, Leiuperinae) in Bahia state, northeastern Brazil, with further data on its advertisement call. Check List, 13(1): 1-4. (PDF)
2016
Carvalho, T.R.; Martins, L.B. & Giaretta, A.A. A new account for the endangered Cerrado Rocket Frog Allobates goianus (Bokermann, 1975) (Anura, Aromobatidae), with comments on taxonomy and conservation. Acta Herpetologica, 11(1): 21-30. (PDF)
Martins, L.B.; Giaretta, A.A.; Carvalho, T.R. & Miwa, R.Y. Vocalizations of Hypsiboas beckeri and H. stenocephalus (Anura: Hylidae), two species of the H. polytaenius group from southeastern Brazil. Phyllomedusa, Journal of Herpetology, 15(1): 51-64. (PDF)
Andrade, M.S.; Ramalho, W.P.; Carvalho, T.R. & Maciel, N.M. The first record of Pseudopaludicola atragula Pansonato, Mudrek, Veiga-Menoncello, Rossa-Feres, Martins & Strüssmann, 2014 (Anura: Leptodactylidae) in the state of Goiás, central Brazil. Check List, 12(5): 1-5. (PDF)
Costa-Campos, C.E.; Carvalho, T.R. & Freire, E.M.X. First record of Pseudopaludicola boliviana Parker, 1927 (Anura, Leptodactylidae, Leiuperinae) in the Brazilian state of Amapá, with comments on its advertisement call and distribution. Check List, 12(6): art. #1991 (PDF)
2015
Carvalho, T.R.; Martins, L.B. & Giaretta, A.A. The complex vocalization of Scinax cardosoi (Anura: Hylidae), with comments on advertisement calls in the S. ruber Clade. Phyllomedusa, Journal of Herpetology, 14(12): 127-137. (PDF)
Carvalho, T.R.; Teixeira, B.F.V.; Duellman, W.E. & Giaretta, A.A. Scinax cruentommus (Anura: Hylidae) in the upper Rio Negro drainage, Amazonas, Brazil, with the re-description of its advertisement call. Phyllomedusa, Journal of Herpetology, 14(2): 139-146. (PDF)
Carvalho, T.R.; Martins, L.B.; Teixeira, B.F.V.; Godinho, L.B. & Giaretta, A.A. Intraspecific variation in acoustic traits and body size, and new distributional records for Pseudopaludicola giarettai Carvalho, 2012 (Anura, Leptodactylidae, Leiuperinae): implications for its congeneric diagnosis. Papéis Avulsos de Zoologia, 55(17): 245-254. (PDF)
Bernardes, C.S.; Carvalho, T.R. & Giaretta, A.A. Advertisement call of Rhinella major (Anura: Bufonidae) from the lower Amazonas River basin with comments on intraspecific variation. Zootaxa, 4012(12): 375-378.
(request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Teixeira, B.F.V.; Martins, L.B. & Giaretta, A.A. Intraspecific variation and new distributional records for Pseudopaludicola species (Anura, Leptodactylidae, Leiuperinae) with trilled advertisement call pattern: diagnostic characters revisited and taxonomic implications. North-Western Journal of Zoology, 11(2): 262-273 (PDF).
2013
Andrade, F.S. & Carvalho, T.R. 2013. A new species of Pseudopaludicola Miranda-Ribeiro (Leiuperinae: Leptodactylidae: Anura) from the Cerrado of southeastern Brazil. Zootaxa, 3608(5): 389-397.
(request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R. & Giaretta, A.A. 2013a. Taxonomic circumscription of Adenomera martinezi (Bokermann, 1956) (Anura: Leptodactylidae: Leptodactylinae), with the recognition of a new cryptic taxon through a bioacoustic approach. Zootaxa, 3701(2): 207-237.
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Carvalho, T.R. & Giaretta, A.A. 2013b. Bioacoustics reveals two new syntopic species of Adenomera Steindachner (Anura: Leptodactylidae: Leptodactylinae) in the Cerrado of central Brazil. Zootaxa, 3731(3): 533-551.
(request pdf and Erratum at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R. & Giaretta, A.A. 2013c. A reappraisal of the geographic distribution of Bokermannohyla sazimai (Cardoso and Andrade, 2012) (Anura: Hylidae) through morphological and bioacoustic approaches. Phyllomedusa, Journal of Herpetology, 12(1): 33-45. (PDF)
Carvalho, T.R.; Giaretta, A.A.; Teixeira, B.F.V. & Martins, L.B. 2013. New bioacoustic and distributional data on Bokermannohyla sapiranga Brandão et al., 2012 (Anura: Hylidae): revisiting its diagnosis in comparison with B. pseudopseudis (Miranda-Ribeiro, 1937). Zootaxa, 3746(2): 383-392.
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Carvalho, T.R.; Leite, F.S.F. & Pezzuti, T.L. 2013. A new species of Leptodactylus Fitzinger (Leptodactylinae, Leptodactylidae, Anura) from rock montane fields of the Chapada Diamantina, northeastern Brazil. Zootaxa, 3701(3): 349-364.
(request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Teixeira, B.F.V.; Martins, L.B. & Giaretta, A.A. 2013. Intraspecific variability of the advertisement call of Chiasmocleis albopunctata (Anura: Microhylidae): note structure as an additional diagnostic character within the genus. Herpetology Notes, 6: 439-446. (PDF)
Carvalho, T.R.; Tolentino, V.C.M. & Giaretta, A.A. 2013. Advertisement call of Rhinella pygmaea (Myers and Carvalho, 1952) (Anura: Bufonidae) from northern State of Rio de Janeiro. Herpetology Notes, 6, 229-231. (PDF)
2012
Carvalho, T.R. 2012. A new species of Pseudopaludicola Miranda-Ribeiro (Leiuperinae: Leptodactylinae: Anura) from the Cerrado of southeastern Brazil with a distinctive advertisement call pattern. Zootaxa, 3328: 47-54.
(request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Facure, K.G. & Giaretta, A.A. 2012. Predation upon eggs of the terrestrial foam nesting frog Leptodactylus fuscus (Leptodactylidae) by larvae of the ground beetle Loxandrus oophagus (Carabidae, Loxandrini). Herpetology Notes, 5: 319-322. (PDF)
Carvalho, T.R.; Giaretta, A.A. & Magrini, L. 2012. A new species of the Bokermannohyla circumdata group (Anura: Hylidae) from southeastern Brazil, with bioacoustic data on seven species of the genus. Zootaxa, 3321: 37-55.
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Carvalho, T.R. & Martins, L.B. 2012. Advertisement call of Haddadus binotatus (Spix, 1824) (Anura: Terrarana: Craugastoridae) from three localities in the State of Rio de Janeiro, with comments on its bioacoustic variability. Herpetology Notes, 5: 419-422. (PDF)
2011
Carvalho, T.R. & Ron, S. 2011. Advertisement call of Leptodactylus labrosus Jiménez de la Espada, 1875 (Anura, Leptodacylidae): an unusual advertisement call within the L. fuscus group. Herpetology Notes, 4: 325-326. (PDF)
Costa, C.; Vanin, S.A. & Carvalho, T.R. 2011. A new Brazilian species of Loxandrus LeConte, 1952, with description of immatures and notes on natural history (Coleoptera: Carabidae: Loxandrini). Zootaxa, 2745, 30-42.
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Costa, H.; Martins, L.B. & Carvalho, T.R. 2011. Xenodon neuwiedii (Serpentes: Xenodontinae): Dieta. Herpetologia Brasileira, 1: 38-40. (PDF)
2010
Carvalho, T.R.; Giaretta, A.A. & Facure, K.G. 2010. A new species of Hypsiboas Wagler (Anura: Hylidae) closely related to H. multifasciatus from southeastern Brazil. Zootaxa, 2521: 37-52.
(request pdf at thiago_decarvalho@yahoo.com.br)
Morais, A.R.; Hepp, F.; Pansonato, A.; Santos, C.E.; Haddad, C.F.B.; Strüssmann, C.; Nascimento, L.B.; Toledo, L.F.; Duarte, M.H.L.; Wachlevski, M.; Simões, P.I.; Bastos, R.P.; Carvalho-e-Silva, S.P.; Dena, S.; Carvalho, T.R.; Forti, L.R. Amphibian sound recordings in Brazil are geographically and taxonomically biased and cover less than two-thirds of native species.
Cassini, C.S.; Carvalho, T.R.; Taucce, P.P.G.; Haddad, C.F.B.; Solé, M. Unexplored urban diversity: a new species of Adenomera (Anura, Leptodactylidae) related to A. ajurauna from the Atlantic Forest of southeastern and southern Brazil.
Zaracho, V.H.; Lavilla, E.O.; Carvalho, T.R.; Motte, M.; Basso, N.G. In pursuit of a nominal anuran species: redescription of Adenomera diptyx (Boettger, 1885) (Leptodactylidae) and description of a closely related new species.
Marinho, P.M.; Santos, M.T.T.; Faivovich, J., Lyra, M.L.; Giaretta, A.A.; Haddad, C.F.B.; Carvalho, T.R. A new species of the Aplastodiscus albosignatus group (Hylinae: Cophomantini) from the northern Mantiqueira Mountain Range.
Accepted for publication
Bang, D.L.; Bockmann, F.A.; Giaretta, A.A.; Carvalho, T.R. The vocal repertoire of two Bokermannohyla (Anura: Hylidae) species, with an overview of the advertisement call diversity in the genus. Journal of Herpetology.
Complete list of publications
2022
Santos, M.T.T.; Barata, I.M.; Ferreira, R.B.; Haddad, C.F.B.; Gridi-Papp, M.; Carvalho, T.R. Complex acoustic signals in Crossodactylodes (Leptodactylidae, Paratelmatobiinae): a frog genus historically regarded as voiceless. Bioacoustics, 31: 175-190. (PDF)
Carvalho, T.R.; Fouquet, A.; Lyra, M.L.; Giaretta, A.A.; Costa-Campos, C.E.; Rodrigues, M.T.; Haddad, C.F.B.; Ron, S.R. Species diversity and systematics of the Leptodactylus melanonotus group (Anura, Leptodactylidae): review of diagnostic traits and a new species from the Eastern Guiana Shield. Systematics and Biodiversity, 20(1): #2089269.
Novaes-e-Fagundes, G.; Lyra, M.L.; Loredam, V.; Carvalho, T.R.; Haddad, C.F.B.; Rodrigues, M.T.; Baldo, D.; Barrasso, D.; Loebmann, D.; Ávila, R.W.; Brusquetti, F.; Prudente, A.L.C.; Wheeler, W.C.; Orrico, V.G.D.; Peloso, P.L.V. A tale of two bellies: systematics of the oval frogs (Anura: Microhylidae: Elachistocleis Parker, 1927). Zoological Journal of the Linnean Society, online first: 1-24.
Taucce, P.P.G.; Costa-Campos, C.E.; Carvalho, T.R.; Michalski, F. Anurans (Amphibia: Anura) of the Brazilian state of Amapá, eastern Amazonia: species diversity and knowledge gaps. European Journal of Taxonomy, 836: 96-130.
Fonseca, W.L.; Almeida, M.R.N.; Nascimento, J.A.F.; Moraes, L.G.L.; Machado, R.A.; Bernarde, P.S.; Costa-Campos, C.E.; Carvalho, T.R. The advertisement call of Allophryne resplendens (Anura: Allophrynidae) from southwestern Brazilian Amazonia. Zootaxa, 3158: 196-200.
2021
Carvalho, T.R.; Seger, K.R.; Magalhães, F.M.; Lourenço, L.B.; Haddad, C.F.B. Systematics and cryptic diversification of Leptodactylus frogs in the Brazilian campo rupestre. Zoologica Scripta, 50(3): 300-317.
Carvalho, T.R.; Moraes, L.C.J.L.; Lima, A.P.; Fouquet, A.; Peloso, P.L.V.; Pavan, D.; Drummond, L.O.; Rodrigues, M.T.; Giaretta, A.A.; Gordo, M.; Neckel-Oliveira, S.; Haddad, C.F.B. Systematics and historical biogeography of Neotropical foam-nesting frogs of the Adenomera heyeri clade (Leptodactylidae), with the description of six new Amazonian species. Zoological Journal of the Linnean Society, 191(2): 395-433. (PDF)
Gazoni, T.*; Lyra, M.L.*; Ron, S.R.; Strüssmann, C.; Baldo, D.; Narimatsu, H.; Pansonato, A.; Schneider R.G.; Giaretta, A.A.; Haddad, Parise-Maltempi, P.P.; Carvalho, T.R*. Revisiting the systematics of the Leptodactylus melanonotus group (Anura: Leptodactylidae): redescription of L. petersii and revalidation of its junior synonyms. Zoologischer Anzeiger, 290: 117-134. (request by e-mail)
* Equal authorship.
Fouquet, A.*; Marinho, P.*; Réjaud, A.; Carvalho, T.R.; Caminer, M.; Jansen, M.; Rainha, R.N.; Rodrigues, M.T.; Werneck, F.P.; Lima, A.P.; Hrbek, T.; Giaretta, A.A.; Venegas, P.J.; Chávez, G; Ron, S. Systematics and biogeography of the Boana albopunctata species group, with the description of two new species from Amazonia. Systematics and Biodiversity, 19(4): 375-399. (request by e-mail)
* Shared co-first authorship.
Carvalho, T.R.; Seger, K.R.; Magalhães, F.M.; Lourenço, L.B.; Haddad, C.F.B. On the availability of the recently proposed nomen Leptodactylus avivoca (Anura, Leptodactylidae). Bionomina, 22(1): 42-45. (request by e-mail)
2020
Carvalho, T.R.; Giaretta, A.A. Diversidade, distribuição e conservação dos anfíbios anuros. In: Vasconcelos, H.L. (Ed.). Ecologia e conservação dos cerrados, campos e florestas do Triângulo Mineiro e sudeste de Goiás. Editora Regência e Arte. (PDF)
Magalhães, F.M.; Lyra, M.L.; Carvalho, T.R.; Baldo, D.; Brusquetti, F.; Burella, P.; Colli, G.R.; Gehara, M.C.; Giaretta, A.A.; Haddad, C.F.B.; Langone, J.A.; López, J.A.; Napoli, M.F.; Santana, D.J.; de Sá, R.O.; Garda, A.A. Taxonomic review of South American butter frogs: phylogeny, biogeographic patterns, and species delimitation in the Leptodactylus latrans species group (Anura: Leptodactylidae). Herpetological Monographs, 34: 131-177. (request by e-mail)
Carvalho, T.R.; Simões, P.I.; Gagliardi-Urrutia, G.; Rojas-Runjaic, F.J.M.; Haddad, C.F.B.; Castroviejo-Fisher, S. A new forest-dwelling frog species of the genus Adenomera (Leptodactylidae) from northwestern Brazilian Amazonia. Copeia, 108(4): 924-937. (request by e-mail)
Taucce, P.P.G.; Costa-Campos, C.E.; Haddad, C.F.B.; Carvalho, T.R. A new Amazonian species of the diminutive frog genus Adelophryne (Anura: Brachycephaloidea: Eleutherodactylidae) from the state of Amapá, northern Brazil. Copeia, 108(4): 746-757. (request by e-mail)
Cassini, C.S.; Taucce, P.P.G.; Carvalho, T.R.; Fouquet, A.; Solé, M.; Haddad, C.F.B.; Garcia, P.C.A. One step beyond a broad molecular phylogenetic analysis: species delimitation of Adenomera marmorata Steindachner, 1867 (Anura: Leptodactylidae). Plos One, 15(2): e0229324. (PDF)
Silva, L.A.; Magalhães, F.M.; Thomassen, H.; Leite, F.S.F.; Garda, A.A.; Brandão, R.A.; Haddad, C.F.B.; Giaretta, A.A.; Carvalho, T.R. Unraveling the species diversity and relationships in the Leptodactylus mystaceus complex (Anura: Leptodactylidae), with the description of three new Brazilian species. Zootaxa, 4779(2): 151-189. (request by e-mail)
Marinho, P.; Costa-Campos, C.E.; Pezzuti, T.L.; Dias-Souza, M.R.; Haddad, C.F.B.; Giaretta, A.A.; Carvalho, T.R. The Amapá tree frog Boana dentei (Bokermann, 1967): diagnosis and redescription (Hylinae: Cophomantini). Journal of Natural History, 54(15-16): 971-990. (request by e-mail)
Carvalho, T.R.*; Moraes, L.J.C.L.*; Angulo, A.; Werneck, F.P.; Icochea, J.; Lima, A.P. New acoustic and molecular data shed light on the poorly known Amazonian frog Adenomera simonstuarti
(Leptodactylidae): implications for distribution and conservation. European Journal of Taxonomy, 682: 1-18. (PDF)
* Shared co-first authorship.
Bang, D.L.; Lisboa, B.S.; Teixeira, B.F.V.; Giaretta, A.A.; Carvalho, T.R. A comparative acoustic analysis of species of Vitreorana (Anura: Centrolenidae) from the Brazilian Atlantic Forest and Cerrado, with a description of the call of V. baliomma and insights into the taxonomic status of Cerrado populations. Phyllomedusa, 19(1): 35-47. (PDF)
Andrade, F.S.; Haga, I.A.; Lyra, M.L.; Carvalho, T.R.; Haddad, C.F.B.; Giaretta, A.A.; Toledo, L.F. Reassessment of the taxonomic status of Pseudopaludicola parnaiba (Anura, Leptodactylidae, Leiuperinae), with the description of a new cryptic species from the Brazilian Cerrado. European Journal of Taxonomy, 679: 1-36. (PDF)
Carvalho, T.R.; Angulo, A.; Barrera, D.A.; Aguilar-Puntriano, C.; Haddad, C.F.B. Hiding in plain sight: a fourth new cryptic species of the Adenomera andreae clade (Anura: Leptodactylidae) from southwestern Amazonia. Herpetologica, 76(3): 304-314. (request by e-mail)
Carvalho, T.R.; Roberto, I.J.; Dias, E.M.; Santos, R.L.; Santos, E.M. The advertisement call of Physalaemus caete (Leptodactylidae), an endangered anuran species endemic to Brazil's northern Atlantic Forest. Zootaxa, 4822(3): 439-442. (request by e-mail)
2019
Carvalho, T.R.; Giaretta, A.A.; Maciel, N.M.; Barrera, D.A.; Aguilar-Puntriano, C.; Haddad, C.F.B.; Kokubum, M.N.C.; Menin, M.; Angulo, A. On the uncertain taxonomic identity of Adenomera hylaedactyla (Cope, 1868) and the composite type series of A. andreae (Müller, 1923) (Anura, Leptodactylidae). Copeia, 107(4): 708-723. (request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Angulo, A.; Kokubum, M.N.C.; Barrera, D.A.; Souza, M.B.; Haddad, C.F.B.; Giaretta, A.A. A new cryptic species of the Adenomera andreae clade from southwestern Amazonia (Anura, Leptodactylidae). Herpetologica, 75(3): 233-246. (request pdf at thiago_decarvalho@yahoo.com.br)
Marinho, P.; Costa-Campos, C.E.; Sanches, P.R.; Carvalho, T.R. The release call of Rhaebo guttatus (Anura: Bufonidae). Studies on Neotropical Fauna and Environment, online first.
(request pdf at thiago_decarvalho@yahoo.com.br)
Santos, M.T.T.; Oliveira, S.H.; Carvalho, T.R.; Zaidan, B.F.; Silva, N.R.; Berneck, B.V.M.; Garcia, P.C.A. A new species of Paratelmatobius (Anura: Leptodactylidae: Paratelmatobiinae) from the Atlantic Forest of southern Brazil. Zootaxa, 4648(3): 473-493. (request pdf at thiago_decarvalho@yahoo.com.br)
Carvalho, T.R.; Haddad, C.F.B.; Gridi-Papp, M. Tonal calls as a bioacoustic novelty in two Atlantic Forest species of Physalaemus (Anura: Leptodactylidae). Acta Herpetologica, 14(1): 21-26. (PDF)
Carvalho, T.R.; Cassini, C.S.; Taucce, P.P.G.; Haddad, C.F.B. A new, morphologically cryptic species of Adenomera closely related to A. araucaria from the Atlantic Forest of southern Brazil (Anura, Leptodactylidae). Journal of Herpetology, 53(2): 131-143.
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Carvalho, T.R.; Giaretta, A.A.; Angulo, A.; Haddad, C.F.B.; Peloso, P.L.V. A new Amazonian species of Adenomera (Anura: Leptodactylidae) from the Brazilian state of Pará: a tody-tyrant voice in a frog. American Museum Novitates, 3919: 1-21. (PDF)
Roberto, I.J.; Santos, E.M.; Carvalho, T.R. The vocalization of Gastrotheca fissipes (Boulenger, 1888) (Anura, Hemiphractidae) from the state of Pernambuco, northeastern Brazil. Zootaxa, 4543(2): 284-286.
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2018
Andrade, F.S.; Haga, I.A.; Lyra, M.L.; Carvalho, T.R.; Haddad, C.F.B., Giaretta, A.A., Toledo, L.F. A new species of Pseudopaludicola (Anura: Leptodactylidae: Leiuperinae) from the state of Minas Gerais, Brazil. European Journal of Taxonomy, 480: 1-25. (PDF)
Marinho, P.; Carvalho, T.R.; Bang, D.L.; Teixeira, B.F.V.; Azarak, P.A.; Costa-Campos, C.E. & Giaretta, A.A. Advertisement calls, intraspecific variation and species diagnosis of six Brazilian species of Elachistocleis (Anura: Microhylidae: Gastrophryninae). Zootaxa, 4521(3): 357-375.
(request pdf at thiago_decarvalho@yahoo.com.br)
Costa-Campos, C.E.; Carvalho, T.R. The advertisement call of the Hoogmoed's harlequin toad Atelopus hoogmoedi Lescure, 1974 from northern Brazil (Anura, Bufonidae). Zootaxa, 4521(1): 141-144.
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Andrade, F.S.; Haga, I.A.; Carvalho, T.R.; Martins, L.B.; Giaretta, A.A. An acoustic and morphometric evaluation of the geographic distribution of Phyllomedusa burmeisteri (Anura: Phyllomedusidae), with comments on P. bahiana. Phyllomedusa, 17(1): 39-59. (PDF)
Vidigal, I.; Carvalho, T.R.; Clemente-Carvalho, R.B.G. & Giaretta, A.A. Vocalizations, tadpole, and natural history of Crossodactylus werneri Pimenta, Cruz & Caramaschi, 2014 (Anura: Hylodidae), with comments on distribution and intraspecific variation. Zootaxa, 4388(1): 061-075.
(request pdf at thiago_decarvalho@yahoo.com.br)
2017
Carvalho, T.R.; Azarak, P.A.; Bang, D.L.; Duellman, W.E. & Giaretta, A.A. A reassessment of the vocalization and distribution of Scinax exiguus (Duellman, 1986) (Anura: Hylidae) in the Amazonian savanna of Roraima, northern Brazil, with the description of its aggressive call. Neotropical Biodiversity, 3(1): 196-202. (PDF)
Carvalho, T.R.; Bang, D.L.; Teixeira, B.F.V. & Giaretta, A.A. First record of Boana alfaroi (Caminer & Ron, 2014) (Anura: Hylidae) in Brazil. Check List, 13(4): 135-139. (PDF)
Haga, I.A.; Carvalho, T.R.; Andrade, F.S. & Giaretta, A.A. Advertisement and aggressive calls of Pithecopus azureus (Anura: Phyllomedusidae) from border of Brazil and Paraguay . Phyllomedusa, 16(1): 47-56. (PDF)
Andrade, F.S.; Carvalho, T.R.; Martins, L.B. & Giaretta, A.A. Reassessment of the vocal repertoire of a nest-building gladiator frog, Boana pardalis (Anura, Hylidae, Cophomantinae): implications for its diagnosis within the B. faber species group. Studies on Neotropical Fauna and Environment, 52(1): 1-11. (PDF)
Bang, D.L.; Carvalho, T.R.; Andrade, F.S.; Aquemi, I.H.; Bernardes, C.S. & Giaretta, A.A. Vocalization of Scinax haddadorum (Anura: Hylidae), with further notes on the vocalization of the morphologically similar S. rupestris. Neotropical Biodiversity, 3(1): 117-124. (PDF)
Andrade, F.S.; Leite, F.S.; Carvalho, T.R.; Bernardes, C.S. & Giaretta, A.A. First record of Pseudopaludicola pocoto Magalhães, Loebmann, Kokubum, Haddad & Garda, 2014 (Anura, Leptodactylidae, Leiuperinae) in Bahia state, northeastern Brazil, with further data on its advertisement call. Check List, 13(1): 1-4. (PDF)
2016
Carvalho, T.R.; Martins, L.B. & Giaretta, A.A. A new account for the endangered Cerrado Rocket Frog Allobates goianus (Bokermann, 1975) (Anura, Aromobatidae), with comments on taxonomy and conservation. Acta Herpetologica, 11(1): 21-30. (PDF)
Martins, L.B.; Giaretta, A.A.; Carvalho, T.R. & Miwa, R.Y. Vocalizations of Hypsiboas beckeri and H. stenocephalus (Anura: Hylidae), two species of the H. polytaenius group from southeastern Brazil. Phyllomedusa, Journal of Herpetology, 15(1): 51-64. (PDF)
Andrade, M.S.; Ramalho, W.P.; Carvalho, T.R. & Maciel, N.M. The first record of Pseudopaludicola atragula Pansonato, Mudrek, Veiga-Menoncello, Rossa-Feres, Martins & Strüssmann, 2014 (Anura: Leptodactylidae) in the state of Goiás, central Brazil. Check List, 12(5): 1-5. (PDF)
Costa-Campos, C.E.; Carvalho, T.R. & Freire, E.M.X. First record of Pseudopaludicola boliviana Parker, 1927 (Anura, Leptodactylidae, Leiuperinae) in the Brazilian state of Amapá, with comments on its advertisement call and distribution. Check List, 12(6): art. #1991 (PDF)
2015
Carvalho, T.R.; Martins, L.B. & Giaretta, A.A. The complex vocalization of Scinax cardosoi (Anura: Hylidae), with comments on advertisement calls in the S. ruber Clade. Phyllomedusa, Journal of Herpetology, 14(12): 127-137. (PDF)
Carvalho, T.R.; Teixeira, B.F.V.; Duellman, W.E. & Giaretta, A.A. Scinax cruentommus (Anura: Hylidae) in the upper Rio Negro drainage, Amazonas, Brazil, with the re-description of its advertisement call. Phyllomedusa, Journal of Herpetology, 14(2): 139-146. (PDF)
Carvalho, T.R.; Martins, L.B.; Teixeira, B.F.V.; Godinho, L.B. & Giaretta, A.A. Intraspecific variation in acoustic traits and body size, and new distributional records for Pseudopaludicola giarettai Carvalho, 2012 (Anura, Leptodactylidae, Leiuperinae): implications for its congeneric diagnosis. Papéis Avulsos de Zoologia, 55(17): 245-254. (PDF)
Bernardes, C.S.; Carvalho, T.R. & Giaretta, A.A. Advertisement call of Rhinella major (Anura: Bufonidae) from the lower Amazonas River basin with comments on intraspecific variation. Zootaxa, 4012(12): 375-378.
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Carvalho, T.R.; Teixeira, B.F.V.; Martins, L.B. & Giaretta, A.A. Intraspecific variation and new distributional records for Pseudopaludicola species (Anura, Leptodactylidae, Leiuperinae) with trilled advertisement call pattern: diagnostic characters revisited and taxonomic implications. North-Western Journal of Zoology, 11(2): 262-273 (PDF).
2013
Andrade, F.S. & Carvalho, T.R. 2013. A new species of Pseudopaludicola Miranda-Ribeiro (Leiuperinae: Leptodactylidae: Anura) from the Cerrado of southeastern Brazil. Zootaxa, 3608(5): 389-397.
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Carvalho, T.R. & Giaretta, A.A. 2013a. Taxonomic circumscription of Adenomera martinezi (Bokermann, 1956) (Anura: Leptodactylidae: Leptodactylinae), with the recognition of a new cryptic taxon through a bioacoustic approach. Zootaxa, 3701(2): 207-237.
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Carvalho, T.R. & Giaretta, A.A. 2013b. Bioacoustics reveals two new syntopic species of Adenomera Steindachner (Anura: Leptodactylidae: Leptodactylinae) in the Cerrado of central Brazil. Zootaxa, 3731(3): 533-551.
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Carvalho, T.R. & Giaretta, A.A. 2013c. A reappraisal of the geographic distribution of Bokermannohyla sazimai (Cardoso and Andrade, 2012) (Anura: Hylidae) through morphological and bioacoustic approaches. Phyllomedusa, Journal of Herpetology, 12(1): 33-45. (PDF)
Carvalho, T.R.; Giaretta, A.A.; Teixeira, B.F.V. & Martins, L.B. 2013. New bioacoustic and distributional data on Bokermannohyla sapiranga Brandão et al., 2012 (Anura: Hylidae): revisiting its diagnosis in comparison with B. pseudopseudis (Miranda-Ribeiro, 1937). Zootaxa, 3746(2): 383-392.
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Carvalho, T.R.; Leite, F.S.F. & Pezzuti, T.L. 2013. A new species of Leptodactylus Fitzinger (Leptodactylinae, Leptodactylidae, Anura) from rock montane fields of the Chapada Diamantina, northeastern Brazil. Zootaxa, 3701(3): 349-364.
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Carvalho, T.R.; Teixeira, B.F.V.; Martins, L.B. & Giaretta, A.A. 2013. Intraspecific variability of the advertisement call of Chiasmocleis albopunctata (Anura: Microhylidae): note structure as an additional diagnostic character within the genus. Herpetology Notes, 6: 439-446. (PDF)
Carvalho, T.R.; Tolentino, V.C.M. & Giaretta, A.A. 2013. Advertisement call of Rhinella pygmaea (Myers and Carvalho, 1952) (Anura: Bufonidae) from northern State of Rio de Janeiro. Herpetology Notes, 6, 229-231. (PDF)
2012
Carvalho, T.R. 2012. A new species of Pseudopaludicola Miranda-Ribeiro (Leiuperinae: Leptodactylinae: Anura) from the Cerrado of southeastern Brazil with a distinctive advertisement call pattern. Zootaxa, 3328: 47-54.
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Carvalho, T.R.; Facure, K.G. & Giaretta, A.A. 2012. Predation upon eggs of the terrestrial foam nesting frog Leptodactylus fuscus (Leptodactylidae) by larvae of the ground beetle Loxandrus oophagus (Carabidae, Loxandrini). Herpetology Notes, 5: 319-322. (PDF)
Carvalho, T.R.; Giaretta, A.A. & Magrini, L. 2012. A new species of the Bokermannohyla circumdata group (Anura: Hylidae) from southeastern Brazil, with bioacoustic data on seven species of the genus. Zootaxa, 3321: 37-55.
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Carvalho, T.R. & Martins, L.B. 2012. Advertisement call of Haddadus binotatus (Spix, 1824) (Anura: Terrarana: Craugastoridae) from three localities in the State of Rio de Janeiro, with comments on its bioacoustic variability. Herpetology Notes, 5: 419-422. (PDF)
2011
Carvalho, T.R. & Ron, S. 2011. Advertisement call of Leptodactylus labrosus Jiménez de la Espada, 1875 (Anura, Leptodacylidae): an unusual advertisement call within the L. fuscus group. Herpetology Notes, 4: 325-326. (PDF)
Costa, C.; Vanin, S.A. & Carvalho, T.R. 2011. A new Brazilian species of Loxandrus LeConte, 1952, with description of immatures and notes on natural history (Coleoptera: Carabidae: Loxandrini). Zootaxa, 2745, 30-42.
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Costa, H.; Martins, L.B. & Carvalho, T.R. 2011. Xenodon neuwiedii (Serpentes: Xenodontinae): Dieta. Herpetologia Brasileira, 1: 38-40. (PDF)
2010
Carvalho, T.R.; Giaretta, A.A. & Facure, K.G. 2010. A new species of Hypsiboas Wagler (Anura: Hylidae) closely related to H. multifasciatus from southeastern Brazil. Zootaxa, 2521: 37-52.
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